In Jasper, Nevada, three young humans, Jack Darby, Miko Nakadai, and Rafael Esquivel, are accidentally caught in the crossfire in a fight between enormous robots that transform into vehicles. They are taken to the base of the heroic Autobots, where their leader Optimus Prime explains that he and his team are from the planet Cybertron. As it turns out, the Autobots had a devastating war against the ruthless Megatron and his followers, the Decepticons long ago on Cybertron. While this is going on, Megatron returns from a three-year-long exile in deep space, bringing with him Dark Energon, a substance able to revive the dead and he sets out to raise an army of the undead and conquer Earth. In the conclusion to Darkness Rising, Megatron is seen in front of the Space Bridge waiting for his army of the undead to come through it; however, the Space Bridge explodes, and Megatron is seen no more.